the WHALE Report
June // 2022
CWR Member News // Published Quarterly

Aerial Observation Study footage of T60, T60F, T60G, T60C, T2B, and T59 was taken during Center for Whale Research Encounter #56, August 23, 2021.
Overhead

or at
Water level
Both perspectives drive the Center for Whale Research’s day-to-day scientific research.
Photographs and videos are taken under Center for Whale Research Federal Permit NMFS #21238.
CWR’s Dave Ellifrit is capturing Orca ID shots from the Center for Whale Research vessel, Orcinus.
QUOTABLE
It is crucial that killer whale research continues in the Salish Sea. While CWR’s observations span many decades, we still have not spanned the entire lifespan of a female killer whale. By monitoring the population’s social structure and demography and applying new data collection and analysis methods, we can continue to expand our understanding of these killer whales [orcas], improve our scientific understanding of animal social evolution, and uncover essential information for population recovery.
Center for Whale Research’s (CWR) Research Director Dr. Michael Weiss’ final paragraph in Why ORCA RESEARCH Matters. He’s explaining why the continued study of Salish Sea orcas is vitally important.
in this issue of
the WHALE Report
QUOTABLE // from the SCIENCE Desk // 2022 SIGHTINGS update
SRKW NEWS // getting to KNOW US
All photographs, videos, and information on WhaleResearch.com are Copyright © 2022 Center for Whale Research.

from the SCIENCE Desk
University of Exeter Ph.D. student Mia Lybkaer Kronborg Nielsen and CWR’s Research Director Dr. Michael Weiss examine UAV/drone video of orcas in CWR’s San Juan Island offices in May 2022. Read Nielsen’s CWR Blog: New evidence of menopause in Bigg’s Transient killer whales.
Why ORCA RESEARCH matters
Center for Whale Research’s new Research Director, Dr. Michael Weiss, explains why the ongoing study of Salish Sea orcas is essential.

CWR photo-identification expert Dave Ellifrit takes a right side ID shot of Bigg’s orca T87 (below) on May 13, 2022, during ORCA SURVEY Encounter #23.

To understand the drivers and consequences of natural processes, scientists need to observe and record that process over relevant timescales. For researchers studying animal populations, this means collecting data on those animals throughout a lifespan or, ideally, over multiple lifespans. When these animals are long-lived, like many large mammals, this can mean several decades of research are needed. That has been the goal of the Center for Whale Research (CWR) for the last 46 years: to understand and conserve the Southern Resident killer whale (SRKW) population through long-term, individual-based scientific research.
Through the ORCA SURVEY program, the primary goal of the Center for Whale Research has been to provide an up-to-date census of the Southern Resident killer whale community. This data is ultimately what scientists and managers use to assess the status and trends of this endangered population over time. The information has been crucial for establishing conservation planning in this population. CWR’s census provided the data to identify the SRKW’s small population size and their decline in the 1990s, ultimately leading to the whales being listed as endangered. This same data revealed key correlations between the Southern Residents’ survival, reproduction, and social cohesion and the abundance of Chinook salmon, which identified salmon recovery as the key conservation action for the SRKWs.
In the last decade, the Center for Whale Research’s long-term data has revealed even more factors driving killer whale survival and identified risk factors in the population. CWR research has shown that even adult killer whales are dependent on their mothers and grandmothers and suffer increased mortality risk when they lose these key family members. Further analysis showed that male killer whales are dependent on their wider social circle: socially integrated males have greater survival probabilities than more peripheral individuals. These social effects, especially the effect of grandmothers and social integration, interact with salmon abundance: social relationships are more important when times are hard. The critical role of social relationships in determining killer whale fitness may have important implications for this population’s trajectories. If the population begins losing mothers and grandmothers during periods when salmon is sparse, we could see a rapid population decline.
While the last several decades have taught us many things about the Southern Residents, there is still much to learn. And what we don’t know may be crucial. While we know that social relationships and prey abundance influence orca survival, we don’t yet fully understand the interplay between ecology, demography, and social structure.
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How do deaths and births change social relationships?
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What are the health consequences for the orcas that remain?
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How does the dependence of individuals on their mothers influence the trajectory of the population?
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What are the long-term effects of reduced social cohesion for the population?
These questions are crucial for fully understanding the state of the SRKW population and setting conservation goals—and will take many more years of data collection and analysis. One of the key tasks right now is to determine which salmon runs have the most influence on killer whale vital rates, which is challenging in a small population with a slow life history. We need to continue the long-term census of the Southern Residents to eventually determine the fine-scale correlates of survival and fecundity in this population.
In addition to continued long-term monitoring, the Center for Whale Research is utilizing new methods to piece together a better picture of killer whale society and its consequences on individuals and populations. Using aerial observations, CWR measures and analyzes fine-scale social interactions, including affiliative social contact, aggression, and prey sharing. This data will be used to determine what drives social relationships and the consequences of particular kinds of social relationships for individual survival and reproduction. Supplementing this, CWR has used intensive image analysis to determine what drives differences in injuries received from conspecifics (“rake marks”) in these killer whales.
While the Southern Residents are the main subjects of CWR’s research, we also conduct regular surveys and observations of Bigg’s [Transient] killer whales. Through these observations, we are beginning to understand the shifting ecology and social structure of this thriving population and better understand their society and life history. This data is both scientifically valuable and important for conservation. It provides a nearly ideal test of many hypotheses about social evolution developed from Resident killer whales. But studying a population of killer whales with many of the same anthropogenic pressures (pollution and noise) but very different ecology and prey availability, CWR and our collaborators can better understand how these factors impact the Southern Residents.
It is crucial that killer whale research continues in the Salish Sea. While CWR’s observations span many decades, we still have not spanned the entire lifespan of a female killer whale. By monitoring the population’s social structure and demography and applying new data collection and analysis methods, we can continue to expand our understanding of these killer whales [orcas], improve our scientific understanding of animal social evolution, and uncover essential information for population recovery.

Dr. Michael Weiss flying CWR’s UAV/drone with Mia Lybkaer Kronborg Nielsen’s assistance during UAV Encounter #2, May 6, 2022.
UAV/drone video footage of the T124s traveling and searching for prey; shortly after this video, they made a Harbor seal kill near Yellow Island (UAV Encounter #5, May 13, 2022)
Center for Whale Research AREAS OF STUDY
The Center for Whale Research’s scientists, staff, and volunteers—with your financial support—have studied the Southern Resident and Bigg’s (Transient) orcas in the Pacific Northwest for FIVE DECADES. In addition, we’ve continuously advocated for the best interests of the gravely endangered Southern Resident orcas.
The CWR team’s commitment to these iconic marine mammals persists today. Our primary goals in 2022 are to continue our essential orca studies while informing decision-makers to make the best choices for the benefit of the Southern Residents: to provide our whales with more wild Chinook salmon in a healthier ocean environment.
ORCA SURVEY
Since 1976, the Center for Whale Research has conducted observation-based studies of killer whales in the Salish Sea. Staff continues to collect detailed demographic data about the Southern Resident killer whale population for the 2022 ORCA SURVEY. This survey includes photo-identification images of members of the SRKW community, observations of births and deaths, and information about the behavior and ecology of the animals, including where the animals are in geographic location and time, their social behavior, and foraging patterns. The dataset provides unprecedented insights into killer whale biology and ecology that can inform management decisions to assist in the recovery of the population.
Aerial Observation Study
The Center for Whale Research-University of Exeter Aerial Observation Study uses drones (i.e., unmanned aerial vehicles/UAVs) to study the Southern Resident and Bigg’s (Transient) killer whales’ social organization and underwater behavior from a new perspective. This study helps better understand the orca ecotypes’ complex lives, revealing factors that influence survival, reproduction, social structure, and the evolution of these species’ unique life history. The study was expanded in 2019, becoming part of a large international project examining the link between social structure and life history in animal populations. CWR Scientific Advisor (Animal Social Networks) and Professor of Animal Behaviour at the University of Exeter, Dr. Darren Croft, summarizes the project: The Evolution of Sex Differences in Mammalian Social Life Histories.
Read about the Center for Whale Research-University of Exeter Aerial Observation Study. The Bigg’s (Transient) killer whale component of the study is done in collaboration with University of Exeter Ph.D. student Mia Nielsen. Read Mia’s July 2021 CWR Blog: New evidence of menopause in Bigg’s Transient killer whales.
RECENT Published Studies
Using drone video footage collected as part of the Center for Whale Research-University of Exeter Aerial Observation Study, CWR’s Dr. Michael Weiss led a team of researchers from the United Kingdom, Italy, and the USA to explore the social lives of orcas. The study, published in the journal Proceedings of the Royal Society B, is entitled: Age and sex influence social interactions, but not associations, within a killer whale pod. The research team included the University of York, University of Washington, and Institute of Biophysics; the Natural Environment Research Council (NERC) provided partial study funding.
Age and sex influence social interactions, but not associations, within a killer whale pod published in May 2021 in the journal Proceedings of the Royal Society B by authors Weiss, M. N., Franks, D. W., Giles, D. A., Youngstrom, S., Wasser, S. K., Balcomb, K. C., Ellifrit, D. K., Domenici, P., Cant, M. A., Ellis, S., Nielsen, M. L. K., Grimes, C., and Croft, D. P. The opening lines of the summary read: “Social structure is a fundamental aspect of animal populations. In order to understand the function and evolution of animal societies, it is important to quantify how individual attributes, such as age and sex, shape social relationships.”
Visit Research Publications at WhaleResearch.com to see a list of other publications where the Center for Whale Research has had involvement, including:
The effect of age, sex, and resource abundance on patterns of rake markings in resident killer whales (Orcinus orca) published in January 2022 in Marine Mammal Science by authors Grimes, C., Brent, L. J. N., Weiss, M. N., Franks, D. W., Balcomb, K. C., Ellifrit, D. K., Ellis, S., and Croft, D. P. The opening lines of the Abstract read: “Fluctuations in aggressive behavior of group-living species can reflect social conflict and competition for resources faced by individuals throughout their lifespan and can negatively impact survival and reproduction. In marine mammals, where social interactions are difficult to observe, tooth rake marks can be used as an indicator of received aggression.”
A long postreproductive life span is a shared trait among genetically distinct killer whale populations published in May 2021 in Ecology and Evolution by authors Nielsen, M. L. K., Ellis, S., Towers, J. R., Doniol-Valcroze, T., Franks, D. W., Cant, M. A., Weiss, M. N., Johnstone, R. A., Balcomb, K. C., Ellifrit, D. K., and Croft, D. P. The opening lines of the Conclusion read: “In conclusion, when taken together with previous work, our findings support the hypothesis that kinship dynamics play a key role in the evolution of a prolonged postreproductive life span in killer whales. However, contrary to our predictions, the timing and expected duration of the postreproductive life span did not vary with the dispersal pattern from the natal group, which likely represents different costs and benefits of helping and harming in the two ecotypes.”
Mixture models as a method for comparative sociality: social networks and demographic change in resident killer whales published in April 2021 in the Ecology and Sociobiology by authors Ellis, S., Franks, D. W., Weiss, M. N., Cant, M. A., Domenici, P., Balcomb, K. C., Ellifrit, D. K., and Croft, D. P.

SIGHTINGS update
2022 Encounter summary
J40 showed her mouthful of teeth and massive tongue
CWR Member DOWNLOAD
For personal use only.
Encounter Location: Haro Strait
Part of Encounter Summary:
J37 was pushing her new calf around and we were able to document that J59 is a female! There were a lot of tail lobs, pec slaps, and several random spyhops. There was also an occasional burst of speed and several tightly packed whales would do a few lunges with much rolling and heavy contact between individuals. We saw one whale get smacked in the face by another’s fluke. At one point, we saw J40 raise her head above the water and open her mouth wide while interacting with other whales. Read all of ORCA SURVEY Encounter #28.
J40 with her mouth open
Encounter #28 with J pod // May 26, 2022
(Photograph by CWR’s Dave Ellifrit).